以下为句子列表：
英文: Up to now, the more comprehensive signaling model of guard cell was developed by Assmann in 99 , which including various principal signaling components and their signaling cascades.
中文: 迄今为止较为完善的气孔保卫细胞信号转导的模式由Assmann (99 )提出，该模式列举了人们已取得共识的信号成分和转导途径，除此之外是否还存在其他新的信号成分及转导途径，仍是人们不断探索的问题。
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