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| This advance enabled a stage 4 follicle to produce hooklets at the ends of barbules that could attach to the grooved barbules of the adjacent barbs and create a pennaceous feather with a closed vane. |
中文意思：
这个进展使得毛囊可以长出末端有小钩的羽小枝，钩住隔壁羽枝的沟状羽小枝，创造出羽片闭合的正羽。 |
以下为相关句子列表(点击查看，可以看对应的解释）：
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