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Only after stage 4 could additional feather variations evolve, including the many specializations we see at stage 5, such as the asymmetrical vane of a flight feather.
中文意思:
一旦演化到阶段4之后,其他各种不同的羽毛才能够演化出来,包括我们在阶段5看到的各种特化羽毛,例如飞羽的不对称羽片。


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Only after drilling and testing is it known whether the trap contains oil or gas. 但准确地讲,他们寻找的是潜在的圈闭,只有在钻井和试油以后才会知道圈闭内是否含油或含气。
Only after liberation did he get a full-time job. 只是在解放后他才找到了专职工作。
Only after programmers began building the code—the instructions a computer uses to execute a program—do they discover the inadequacy of their designs. 程式设计师只有在开始撰写程式码(电脑用来执行软体的指令)之后,才会发现自身设计的缺失。
Only after receiving the detailed report, can we work out the way to handle the problem. 只有接到详细报告以后我们才能找出对这一问题的处理办法。
Only after several months did I see the results of my work. 过了几个月,我才看到工作的结果。
Only after stage 4 could additional feather variations evolve, including the many specializations we see at stage 5, such as the asymmetrical vane of a flight feather. 一旦演化到阶段4之后,其他各种不同的羽毛才能够演化出来,包括我们在阶段5看到的各种特化羽毛,例如飞羽的不对称羽片。
Only after ten years in the country did she gain her citizenship. 她在这个国家住了十年后才取得了公民身份。
Only after that we had phone in the dorm, will my parents call me up every month. 而我却总是在忙着玩各种新出的游戏,有心无心的搭两句。
Only after the mother comes to take charge8), can the father penguin go to eat and rest. 只有当帝企鹅妈妈接手育雏工作之后,帝企鹅爸爸才能去进食休息。
Only after the next economic downturn will it become clear whether the tighter rules have made American capitalism stronger or weaker. 只有在下一轮的经济衰退期结束后,严苛的制度是促使了还是削弱了美国的资本主义经济才会情势明朗。
Only after the staggering extended battle sequence does Spielberg introduce you to the characters and story. 在这一长段令人震惊的战争场景之后,斯皮尔伯格才将影片中的人物和故事展现在观众面前。
 
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